Search Results

Overview

Uniprot IDP63017
Protein NameHeat shock cognate 71 kDa protein
Gene NameHspa8
OrganismMus musculus

Kla Sites from experimental identification

Position Flanking peptide
108 PKVQVEYKGETKSFY
500 KSTGKENKITITNDK
507 KITITNDKGRLSKED
512 NDKGRLSKEDIERMV
601 HQQKELEKVCNPIIT
71 TNTVFDAKRLIGRRF

Function

Molecular chaperone implicated in a wide variety of cellular processes, including protection of the proteome from stress, folding and transport of newly synthesized polypeptides, chaperone-mediated autophagy, activation of proteolysis of misfolded proteins, formation and dissociation of protein complexes, and antigen presentation (PubMed:30718432). Plays a pivotal role in the protein quality control system, ensuring the correct folding of proteins, the re-folding of misfolded proteins and controlling the targeting of proteins for subsequent degradation. This is achieved through cycles of ATP binding, ATP hydrolysis and ADP release, mediated by co-chaperones. The co-chaperones have been shown to not only regulate different steps of the ATPase cycle of HSP70, but they also have an individual specificity such that one co-chaperone may promote folding of a substrate while another may promote degradation. The affinity of HSP70 for polypeptides is regulated by its nucleotide bound state. In the ATP-bound form, it has a low affinity for substrate proteins. However, upon hydrolysis of the ATP to ADP, it undergoes a conformational change that increases its affinity for substrate proteins. HSP70 goes through repeated cycles of ATP hydrolysis and nucleotide exchange, which permits cycles of substrate binding and release. The HSP70-associated co-chaperones are of three types: J-domain co-chaperones HSP40s (stimulate ATPase hydrolysis by HSP70), the nucleotide exchange factors (NEF) such as BAG1/2/3 (facilitate conversion of HSP70 from the ADP-bound to the ATP-bound state thereby promoting substrate release), and the TPR domain chaperones such as HOPX and STUB1. Plays a critical role in mitochondrial import, delivers preproteins to the mitochondrial import receptor TOMM70. Acts as a repressor of transcriptional activation. Inhibits the transcriptional coactivator activity of CITED1 on Smad-mediated transcription. Component of the PRP19-CDC5L complex that forms an integral part of the spliceosome and is required for activating pre-mRNA splicing. May have a scaffolding role in the spliceosome assembly as it contacts all other components of the core complex. Binds bacterial lipopolysaccharide (LPS) and mediates LPS-induced inflammatory response, including TNF secretion by monocytes. Substrate recognition component in chaperone-mediated autophagy (CMA), a selective protein degradation process that mediates degradation of proteins with a -KFERQ motif: HSPA8/HSC70 specifically recognizes and binds cytosolic proteins bearing a -KFERQ motif and promotes their recruitment to the surface of the lysosome where they bind to lysosomal protein LAMP2 (PubMed:30718432). KFERQ motif-containing proteins are eventually transported into the lysosomal lumen where they are degraded (PubMed:30718432). In conjunction with LAMP2, facilitates MHC class II presentation of cytoplasmic antigens by guiding antigens to the lysosomal membrane for interaction with LAMP2 which then elicits MHC class II presentation of peptides to the cell membrane. Participates in the ER-associated degradation (ERAD) quality control pathway in conjunction with J domain-containing co-chaperones and the E3 ligase STUB1. It is recruited to clathrin-coated vesicles through its interaction with DNAJC6 leading to activation of HSPA8/HSC70 ATPase activity and therefore uncoating of clathrin-coated vesicles (By similarity)

Protein Sequence

10 MSKGPAVGID 20 LGTTYSCVGV 30 FQHGKVEIIA 40 NDQGNRTTPS 50 YVAFTDTERL 60 IGDAAKNQVA 70 MNPTNTVFDA 80 KRLIGRRFDD 90 AVVQSDMKHW 100 PFMVVNDAGR 110 PKVQVEYKGE 120 TKSFYPEEVS 130 SMVLTKMKEI 140 AEAYLGKTVT 150 NAVVTVPAYF 160 NDSQRQATKD 170 AGTIAGLNVL 180 RIINEPTAAA 190 IAYGLDKKVG 200 AERNVLIFDL 210 GGGTFDVSIL 220 TIEDGIFEVK 230 STAGDTHLGG 240 EDFDNRMVNH 250 FIAEFKRKHK 260 KDISENKRAV 270 RRLRTACERA 280 KRTLSSSTQA 290 SIEIDSLYEG 300 IDFYTSITRA 310 RFEELNADLF 320 RGTLDPVEKA 330 LRDAKLDKSQ 340 IHDIVLVGGS 350 TRIPKIQKLL 360 QDFFNGKELN 370 KSINPDEAVA 380 YGAAVQAAIL 390 SGDKSENVQD 400 LLLLDVTPLS 410 LGIETAGGVM 420 TVLIKRNTTI 430 PTKQTQTFTT 440 YSDNQPGVLI 450 QVYEGERAMT 460 KDNNLLGKFE 470 LTGIPPAPRG 480 VPQIEVTFDI 490 DANGILNVSA 500 VDKSTGKENK 510 ITITNDKGRL 520 SKEDIERMVQ 530 EAEKYKAEDE 540 KQRDKVSSKN 550 SLESYAFNMK 560 ATVEDEKLQG 570 KINDEDKQKI 580 LDKCNEIISW 590 LDKNQTAEKE 600 EFEHQQKELE 610 KVCNPIITKL 620 YQSAGGMPGG 630 MPGGFPGGGA 640 PPSGGASSGP TIEEVD

Gene Ontology

Classification GO ID Description
Cellular Component GO:0032279 asymmetric synapse
Molecular Function GO:0005102 signaling receptor binding
Molecular Function GO:0031625 ubiquitin protein ligase binding
Molecular Function GO:0051082 unfolded protein binding
Biological Process GO:0046034 ATP metabolic process
Biological Process GO:0071276 cellular response to cadmium ion
Biological Process GO:0034605 cellular response to heat
Biological Process GO:0070301 cellular response to hydrogen peroxide
Biological Process GO:0021549 cerebellum development
Biological Process GO:0061684 chaperone-mediated autophagy
Biological Process GO:1904764 chaperone-mediated autophagy translocation complex disassembly
Biological Process GO:0072318 clathrin coat disassembly
Biological Process GO:0006351 DNA-templated transcription
Biological Process GO:0044849 estrous cycle
Biological Process GO:0030900 forebrain development
Biological Process GO:0000082 G1/S transition of mitotic cell cycle
Biological Process GO:0044829 host-mediated activation of viral genome replication
Biological Process GO:0044788 host-mediated perturbation of viral process
Biological Process GO:0001822 kidney development
Biological Process GO:0061738 late endosomal microautophagy
Biological Process GO:0098880 maintenance of postsynaptic specialization structure
Biological Process GO:0000398 mRNA splicing, via spliceosome
Biological Process GO:0010667 negative regulation of cardiac muscle cell apoptotic process
Biological Process GO:0045892 negative regulation of DNA-templated transcription
Biological Process GO:1900226 negative regulation of NLRP3 inflammasome complex assembly
Biological Process GO:1902904 negative regulation of supramolecular fiber organization
Biological Process GO:0043085 positive regulation of catalytic activity
Biological Process GO:0030335 positive regulation of cell migration
Biological Process GO:0160020 positive regulation of ferroptosis
Biological Process GO:0010628 positive regulation of gene expression
Biological Process GO:0097214 positive regulation of lysosomal membrane permeability
Biological Process GO:0048026 positive regulation of mRNA splicing, via spliceosome
Biological Process GO:0050766 positive regulation of phagocytosis
Biological Process GO:1904592 positive regulation of protein refolding
Biological Process GO:0045862 positive regulation of proteolysis
Biological Process GO:0001916 positive regulation of T cell mediated cytotoxicity
Biological Process GO:0046777 protein autophosphorylation
Biological Process GO:0030163 protein catabolic process
Biological Process GO:0006457 protein folding
Biological Process GO:0006606 protein import into nucleus
Biological Process GO:0042026 protein refolding
Biological Process GO:0061740 protein targeting to lysosome involved in chaperone-mediated autophagy
Biological Process GO:0044743 protein transmembrane import into intracellular organelle
Biological Process GO:0032984 protein-containing complex disassembly
Biological Process GO:0051726 regulation of cell cycle
Biological Process GO:0099175 regulation of postsynapse organization
Biological Process GO:0061635 regulation of protein complex stability
Biological Process GO:0031647 regulation of protein stability
Biological Process GO:0014823 response to activity
Biological Process GO:0032355 response to estradiol
Biological Process GO:0045471 response to ethanol
Biological Process GO:0010045 response to nickel cation
Biological Process GO:1990834 response to odorant
Biological Process GO:0032570 response to progesterone
Biological Process GO:0042594 response to starvation
Biological Process GO:0009410 response to xenobiotic stimulus
Biological Process GO:0007519 skeletal muscle tissue development
Biological Process GO:1990832 slow axonal transport
Cellular Component GO:0005776 autophagosome
Cellular Component GO:0030424 axon
Cellular Component GO:0009986 cell surface
Cellular Component GO:0005737 cytoplasm
Cellular Component GO:0005829 cytosol
Cellular Component GO:0030425 dendrite
Cellular Component GO:0043198 dendritic shaft
Cellular Component GO:0043197 dendritic spine
Cellular Component GO:0070062 extracellular exosome
Cellular Component GO:0005615 extracellular space
Cellular Component GO:0098978 glutamatergic synapse
Cellular Component GO:0098690 glycinergic synapse
Cellular Component GO:0005882 intermediate filament
Cellular Component GO:0005770 late endosome
Cellular Component GO:0031906 late endosome lumen
Cellular Component GO:1990836 lysosomal matrix
Cellular Component GO:0005765 lysosomal membrane
Cellular Component GO:0042470 melanosome
Cellular Component GO:0005874 microtubule
Cellular Component GO:0043209 myelin sheath
Cellular Component GO:0044309 neuron spine
Cellular Component GO:0043025 neuronal cell body
Cellular Component GO:0005730 nucleolus
Cellular Component GO:0005634 nucleus
Cellular Component GO:0043204 perikaryon
Cellular Component GO:0048471 perinuclear region of cytoplasm
Cellular Component GO:0001917 photoreceptor inner segment
Cellular Component GO:0098684 photoreceptor ribbon synapse
Cellular Component GO:0005886 plasma membrane
Cellular Component GO:0098794 postsynapse
Cellular Component GO:0014069 postsynaptic density
Cellular Component GO:0099634 postsynaptic specialization membrane
Cellular Component GO:0098793 presynapse
Cellular Component GO:0101031 protein folding chaperone complex
Cellular Component GO:0032991 protein-containing complex
Cellular Component GO:0000974 Prp19 complex
Cellular Component GO:1990904 ribonucleoprotein complex
Cellular Component GO:0005681 spliceosomal complex
Cellular Component GO:0008021 synaptic vesicle
Cellular Component GO:0043195 terminal bouton
Molecular Function GO:0031686 A1 adenosine receptor binding
Molecular Function GO:0005524 ATP binding
Molecular Function GO:0016887 ATP hydrolysis activity
Molecular Function GO:0140545 ATP-dependent protein disaggregase activity
Molecular Function GO:0055131 C3HC4-type RING finger domain binding
Molecular Function GO:1990833 clathrin-uncoating ATPase activity
Molecular Function GO:0019899 enzyme binding
Molecular Function GO:0001664 G protein-coupled receptor binding
Molecular Function GO:0031072 heat shock protein binding
Molecular Function GO:0042277 peptide binding
Molecular Function GO:0001786 phosphatidylserine binding
Molecular Function GO:1904593 prostaglandin binding
Molecular Function GO:0044183 protein folding chaperone
Molecular Function GO:0051087 protein-folding chaperone binding
Molecular Function GO:0030674 protein-macromolecule adaptor activity
Molecular Function GO:0048018 receptor ligand activity
Molecular Function GO:0003723 RNA binding

Reference

[1] Chang J, Wu W, Qian P, Lu Z, He X et al.. Multi-omics study on the effect of moderate-intensity exercise on protein lactylation in mouse muscle tissue.. Front Cell Dev Biol 12:1472338. 2024. PMID: 39935788.

[2] Wu D, Tang Y, Li X, Xiong S, Zhang Z et al.. Characterization of protein lactylation in healthy and ischemic mouse hearts.. Front Cardiovasc Med 12:1644886. 2025. PMID: 41089239.