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Overview

Uniprot IDQ01768
Protein NameNucleoside diphosphate kinase B
Gene NameNme2
OrganismMus musculus

Kla Sites from experimental identification

Position Flanking peptide
49 RASEEHLKQHYIDLK

Function

Catalyzes the transfer of a gamma-phosphoryl group from a nucleoside triphosphate, mainly ATP, to a nucleoside diphosphate via a ping-pong mechanism involving a phosphohistidine intermediate, therefore contributing to the nucleoside triphosphate homeostasis (PubMed:1325378). Also functions as a histidine protein kinase by transferring the phosphoryl group from the phosphohistidine intermediate to a histidine residue in target proteins (By similarity). Phosphorylates the GNB1 subunit of heterotrimeric G proteins at 'His-266', generating a high-energy phosphate group that promotes GTP formation and enables receptor-independent activation of heterotrimeric G proteins (By similarity). Also phosphorylates KCNN4 at 'His-358', leading to activation of its intermediate conductance calcium-activated potassium channel activity, Ca(2+) influx, and subsequent activation of B and T cells. Additionally involved in transcriptional regulation through direct DNA binding and chromatin remodeling. In this context, functions as a single-stranded DNA binding protein that binds and stabilizes the G-quadruplex (G4) structures within the nuclease hypersensitive element (NHE) III(1) region of the MYC gene promoter, facilitating recruitment of additional single-strand DNA binding proteins and activation of MYC transcription. G4 DNA-binding activity is independent of its nucleoside diphosphate kinase function and recognizes both folded and unfolded G4 structures (By similarity). With NME1, may regulate acetyl-CoA (AcCoA) usage between histone acetylation and fatty acid synthesis by targeting AcCoA release at ATP-rich, HAT-associated chromatin regions (PubMed:37672589). Also negatively regulates Rho activity by interacting with AKAP13/LBC (By similarity)

Protein Sequence

10 MANLERTFIA 20 IKPDGVQRGL 30 VGEIIKRFEQ 40 KGFRLVAMKF 50 LRASEEHLKQ 60 HYIDLKDRPF 70 FPGLVKYMNS 80 GPVVAMVWEG 90 LNVVKTGRVM 100 LGETNPADSK 110 PGTIRGDFCI 120 QVGRNIIHGS 130 DSVESAEKEI 140 HLWFKPEELI 150 DYKSCAHDWV YE

Gene Ontology

Classification GO ID Description
Cellular Component GO:0071944 cell periphery
Cellular Component GO:0005737 cytoplasm
Cellular Component GO:0005829 cytosol
Cellular Component GO:0005882 intermediate filament
Cellular Component GO:0030027 lamellipodium
Cellular Component GO:0031966 mitochondrial membrane
Cellular Component GO:0005739 mitochondrion
Cellular Component GO:0043209 myelin sheath
Cellular Component GO:0005634 nucleus
Cellular Component GO:0048471 perinuclear region of cytoplasm
Cellular Component GO:0001726 ruffle
Molecular Function GO:0005524 ATP binding
Molecular Function GO:0120225 coenzyme A binding
Molecular Function GO:0003677 DNA binding
Molecular Function GO:0001216 DNA-binding transcription activator activity
Molecular Function GO:0019899 enzyme binding
Molecular Function GO:0005504 fatty acid binding
Molecular Function GO:0051880 G-quadruplex DNA binding
Molecular Function GO:0019003 GDP binding
Molecular Function GO:1901363 heterocyclic compound binding
Molecular Function GO:0042802 identical protein binding
Molecular Function GO:0019215 intermediate filament binding
Molecular Function GO:0016301 kinase activity
Molecular Function GO:0046872 metal ion binding
Molecular Function GO:0004550 nucleoside diphosphate kinase activity
Molecular Function GO:0004673 protein histidine kinase activity
Molecular Function GO:0004674 protein serine/threonine kinase activity
Biological Process GO:0006241 CTP biosynthetic process
Biological Process GO:0006351 DNA-templated transcription
Biological Process GO:0006231 dTMP biosynthetic process
Biological Process GO:0006183 GTP biosynthetic process
Biological Process GO:0007229 integrin-mediated signaling pathway
Biological Process GO:0046042 ITP biosynthetic process
Biological Process GO:0043066 negative regulation of apoptotic process
Biological Process GO:0009142 nucleoside triphosphate biosynthetic process
Biological Process GO:0045893 positive regulation of DNA-templated transcription
Biological Process GO:2000144 positive regulation of DNA-templated transcription initiation
Biological Process GO:0050679 positive regulation of epithelial cell proliferation
Biological Process GO:0045618 positive regulation of keratinocyte differentiation
Biological Process GO:0045944 positive regulation of transcription by RNA polymerase II
Biological Process GO:0042981 regulation of apoptotic process
Biological Process GO:0042304 regulation of fatty acid biosynthetic process
Biological Process GO:0006228 UTP biosynthetic process

Reference

[1] Chang J, Wu W, Qian P, Lu Z, He X et al.. Multi-omics study on the effect of moderate-intensity exercise on protein lactylation in mouse muscle tissue.. Front Cell Dev Biol 12:1472338. 2024. PMID: 39935788.

[2] Wu D, Tang Y, Li X, Xiong S, Zhang Z et al.. Characterization of protein lactylation in healthy and ischemic mouse hearts.. Front Cardiovasc Med 12:1644886. 2025. PMID: 41089239.